
Courses for PhD students
Postgraduate education
Continuous education is an essential part of PhD studies, and students can take a wide range of courses in agreement with their supervisor and director of studies. The number of course credits required for a licentiate or doctoral degree is determined by the study plans for each subject; the courses that the students must follow are decided in the individual study plan.
Mandatory courses
There are three mendatory courses for all PhD students in biology.
- An introductory course to studying at postgraduate level, to take during the first year (3hp)
- A course in research ethics of at least 2 higher education credits (2hp)
- If you teach at basic and advanced level during your PhD studies, you also have to take a course in university education: Academic teacher training course (5hp)
Other courses
Courses offered by the Biology Section:
There are a number of PhD courses offered at both the Evolutionary Biology Center (EBC) and the Biomedical Center (BMC) by the three research departments within the Biology Section.
More courses, at Uppsala and elsewhere:
SciLifeLab offers many popular courses among PhD students, centered around wet lab methods and bioinformatics. Check regularly for updates.
There are a number of courses offered by the Faculty of Science and Technology at Uppsala University.
Some other opportunities:
Workshops in genomics - Cesky Krumlov
Graduate Research School in Genomic Ecology - Lund University
EMBO courses
Advanced Research Training Courses - Marine Biological Laboratory at Woods Hole
Cold Spring Harbour Laboratory courses
For suggesting topics, get in touch with the PhD School director (mario.vallejo-marin@ebc.uu.se).

More on Postgraduate Studies
All the information you need about postgraduate studies can be found at the faculty web pages, both for enrolled and future students.
Director of studies for PhD students

Department of Cell and Molecular Biology
Anders.Virtanen@icm.uu.se
018-471 4908
Department of Ecology and Genetics
Peter.Eklov@ebc.uu.se
018-471 2720
Department of Organismal Biology
carina.schlebusch@ebc.uu.se
+46 76 3063341